Adnan el Shukrijumah was killed in a raid in north-western Pakistan, near the Afghan border, the military said.
The FBI describes him as al-Qaeda's global operations chief, a post once held by the alleged mastermind of the 9/11 attacks, Khalid Sheikh Mohammed.
Shukrijumah was born in Saudi Arabia and lived for several years in the US.
He was named in a US federal indictment as a conspirator in the case against three men accused of plotting suicide bomb attacks on New York's subway system in 2009.
He is also suspected of having played a role in plotting al-Qaeda attacks in Panama, Norway and the UK.
Saturday's raid took place in the Shinwarsak region of South Waziristan, bordering Afghanistan.
The region is a base for the Pakistani Taliban and its allies.
Pakistan's military said two other militants were killed in the raid. One soldier was also reportedly killed, and another was hurt.
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